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People
Name:
 ZHAO Fangchen
Education:
 Ph.D.
Research direction:
  
Academic
title:
 Professor
Postal Code:
 210008
Subject
categories:
 palaeontology
Mailing
Address:
 39 East Beijing Road, Nanjing, 210008, China
E-mail:
 fczhao@nigpas.ac.cn

Resume:
Education and academic degrees
Sept., 1999-Jul., 2003, China University of Geoscience (Beijing). Major in Geology. Bachelor of Science.
Sept., 2003-Jul., 2006, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences. Major in Stratigraphy and Palaeontology. Master of Science.
Sept., 2006-Jul., 2009, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences. Major in Stratigraphy and Palaeontology. Doctor of Science.
Professional occupation
2009.8-201.12: Assistant Professor in Geology and Palaeontology of Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences
2010.12-2011.5: Visiting scholar, Royal Ontario Museum, Canada
2012.7-2013.7 : Visiting scholar, University of Southern California, USA 
2013.1-2017.3 : Associate Professor in Geology and Palaeontology of Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences  
2017.3-       : Professor in Geology and Palaeontology of Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences  
Awards and Honors
Excellent Master's thesis of the Jiangsu Province, 2007
“outstanding graduate of Graduate University of Chinese Academy of Sciences”, 2009
 “Best student Poster Presentation” on the occasion of the 2009 International Conference on the Cambrian explosion (Banff, Canada)
The member of the Youth Innovation Promotion Association，Chinese Academy Sciences, 2014-
Research topics 
My personal research interests mainly focus on the study of the first complex marine communities from the period just following the Early Cambrian. The “Cambrian explosion” represents the sudden worldwide appearance and rapid diversification of most major animal groups, starting about 540 million years ago. The record of this critical event is preserved in a series of exceptional fossil deposits (Konservat-Lagerst？tten), of which the early Cambrian Chengjiang fauna from China is certainly the most famous. My research is currently oriented in four main areas on the Chenjiang fauna:
1) to elucidate the ecology and affinities of dozens of specific fossil taxa in order to reveal the origin and relationship of major animal body plans and the acquisition of functional novelties during the “Cambrian explosion”;
2) to disclose the interaction between environmental signals and community composition in Cambrian biota;
3) to better understand community patterns at varying temporal and spatial scales, especially between sites from different paleogeographic and paleoecologic settings.
4) to better understand the taphonomy of extraordinary fossils.
Direction:
Achievements:
Publications:

1.        Sun, H.J., Zhao, F.C., Steiner, M., Li, G.X., Na, L., Pan, B., Yin, Z.J., Zeng, H., Van Iten, H., Zhu, M.Y.*, 2020. Skeletal faunas of the lower Cambrian Yu’anshan Formation, eastern Yunnan, China: metazoan diversity and community structure during the Cambrian Age 3. Palaeogeography, Palaeoclimatology, Palaeoecology, 542(2020): 109580. 

2.        Aria, C., Zhao, F.C., Zeng, H., Guo, J., Zhu, M.Y.*, 2020. Fossils from South China redefine the ancestral euarthropod body plan. BMC Evolutionary Biology, https://doi.org/10.1186/s12862-019-1560-7. 20:4. 

3.        Luo, C. *, Zhao, F.C., Zeng, H., 2020. The first report of a vauxiid sponge from the Cambrian Chengjiang Biota. Journal of Paleontology, 94(1):28-33. doi: 10.1017/jpa.2019.52 

4.        Sun, H.J., Yin, Z.J.*, Li, G.X., Zhao, F.C., Zeng, H., Zhu, M.Y., 2019. Periodic shell decollation as an ecology‐driven strategy in the early Cambrian Cupitheca. Palaeontology, https://doi.org/10.1111/pala.12463. 

5.        朱茂炎*, 赵方臣, 殷宗军, 曾晗, 李国祥, 2019. 中国的寒武纪大爆发研究: 进展与展望. 中国科学：地球科学，49(10): 1455-1490. 

6.        Sun, Z.X., Zeng, H., and Zhao, F.C.* 2019. First occurrence of the Cambrian arthropod Sidneyia Walcott, 1911 outside of Laurentia. Geological Magazine, https://doi.org/10.1017/ S0016756819000864. 

7.        Bottjer, D.J.*, Yin, Z.J., Zhao, F.C., Zhu, M.Y. 2019. Comparative taphonomy and phylogenetic signal of phosphatized Weng’an and Kuanchuanpu Biotas. Precambrian Research, https://doi.org/10.1016/j.precamres.2019.105408 

8.        Zhu, M.Y.*, Yang, A.H., Yuan, J.L., Li, G.X., Zhang, J.M., Zhao, F.C., Ahn, S.-Y., Miao, L.Y. 2019. Cambrian integrative stratigraphy and timescale of China. SCIENCE CHINA Earth Sciences, 62(1): 25-60. doi.org/10.1007/s11430-017-9291-0. [朱茂炎*，杨爱华，袁金良，李国祥，张俊明，赵方臣，Ahn Soo-Yeun，苗兰云. 2019. 中国寒武纪综合地层和时间框架. 中国科学：地球科学，49(1): 26-65. doi.org/10.1360/N072017-00370] 

9.        孙智新, 王平丽, 赵方臣, 袁金良*. 2018. 东平寒武系馒头组顶部胸针球接子(Peronopsis Hawle and Corda，1847)的新材料. 古生物学报，57（3）：287-294. 

10.        胡春林*, 赵方臣, 纪秋梅, 朱茂炎. 2018. 宁镇山脉的盖帽碳酸盐岩. 地层学杂志，42(4): 381-392. 

11.        董存志, 胡世学*, 赵方臣. 2018. 葫芦虫化石在寒武纪关山生物群中的发现及其意义. 古生物学报，57(4): 424-428 . 

12.        Sun, H.J.*, Zhao, F.C., Wen, R.Q., Zeng, H., Peng, J. 2018. Feeding strategy and locomotion of Cambrian hyolithides. Palaeoworld, 27(3): 334-342. 

13.        Sun, H.J., Smith, M.R.*, Zeng, H., Zhao, F.C.*, Li, G.X., Zhu, M.Y. 2018. Hyoliths with pedicles illuminate the origin of the brachiopod body plan. Proc. R. Soc. B., 285：20181780. DOI:10.1098/rspb.2018.1780 

14.        Yin, Z.J.*, Zhao, D.D., Pan, B., Zhao, F.C., Zeng, H., Li, G.X., Bottjer, D., Zhu, M.Y. 2018. Early Cambrian animal diapause embryos revealed by X-ray tomography. Geology, 46(5):387-390. https://doi.org/10.1130/G40081.1 （cover article） 

15.        Zeng, H., Zhao, F.C.*, Yin, Z.J., Zhu, M.Y. 2018. A new radiodontan oral cone with a unique combination of anatomical features from the early Cambrian Guanshan Lagerst?tte, eastern Yunnan, South China. Journal of Paleontology，92（1）：40-48. https://doi.org/10.1017/jpa.2017.77. 

16.        Zeng, H., Zhao, F.C.*, Yin, Z.J., Zhu, M.Y. 2018. Morphology of diverse radiodontan head sclerites from the early Cambrian Chengjiang Lagerst?tte, southwest China. Journal of Systematic Palaeontology, 16(1): 1-37. doi: 10.1080/14772019.2016.1263685. 

17.        Hu, S.X. *, Erdtmann, B.D., Steiner, M., Zhang, Y.D., Zhao, F.C., Zhang, Z.F., Zhang, Z.L., Han, J. 2018. Malongitubus: a possible pterobranch hemichordate from the early Cambrian of South China. Journal of Paleontology，92（1）：26-32. doi: 10.1017/jpa.2017.134. 

18.        Zhao, F.C.*, Smith Martin R.*, Yin, Z.J., Zeng, H., Li, G.X., Zhu, M.Y. 2017. Orthrozanclus elongata n. sp. and the significance of sclerite-covered taxa for early trochozoan evolution. Scientific Reports 7, Article number: 16232. doi:10.1038/s41598-017-16304-6. 

19.        Zhao, F.C.*, Guo, J., Yin, Z.J., Li, G.X., Zhu, M.Y. 2017. The extraordinarily preserved Cambrian Chengjiang Biota and the Ediacaran-Cambrian Boundary section at Meishucun, Yunnan. In: Tong, J.N. and Wang, Y.D. (eds). Linking the land and the sea-Field guide book for the 2nd Joint Conference of the Palaeontological Society of China and the Pal?ontologische Gesellschaft. University of Science and Technology of China Press. 113–173. 

20.        Zeng, H., Zhao, F.C.*, Yin, Z.J., Zhu, M.Y. 2017. Appendages of an early Cambrian metadoxidid trilobite from Yunnan, SW China support mandibulate affinities of trilobites and artiopods. Geological Magazine，154（6）：1306-1328. 

21.        Hu, S.X.*, Zhu, M.Y., Zhao, F.C., Steiner, M. 2017. A crown group priapulid from the early Cambrian Guanshan Lagerst?tte. Geological Magazine，154（6）：1329-1333. 

22.        Zhu, M.Y.*, Zhuravlev, A.Yu., Wood, R.A., Zhao, F.C., Sukhov, S.S., 2017. A deep root for the Cambrian explosion: Implications of new bio- and chemostratigraphy from the Siberian Platform. Geology, 45(5): 459-462. doi:10.1130/G38865.1. 

23.        Wood, R.A.*, Zhuravlev, A.Yu., Sukhov, S.S., Zhu, M., Zhao, F. 2017. Demise of Ediacaran dolomitic seas marks widespread biomineralization on the Siberian Platform. Geology, 45(1):27-30. doi:10.1130/G38367.1 

24.        Yin, Z.J.*, Zhu, M.Y., Bottjer, D.J., Zhao, F.C., Tafforeau, P. 2016. Meroblastic cleavage identifies some Ediacaran Doushantuo (China) embryo-like fossils as metazoans. Geology, 44(9):735-738.doi:10.1130/G38262.1 

25.        朱茂炎，张俊明，杨爱华，李国祥，赵方臣，吕苗，殷宗军，2016. 华南新元古代地层、生-储-盖层发育与沉积环境，见: 孙杻、王铁冠 编. 中国东部中-新元古界地质学与油气资源. 北京：科学出版社，107-135. 

26.        Zhu, S.X., Zhu, M.Y.*, Knoll, A.H., Yin, Z.J., Zhao, F.C., Sun, S., Qu, Y., Shi, M., Liu, H. 2016. Decimetre-scale multicellular eukaryotes from the 1.56-billion-year-old Gaoyuzhuang Formation in North China. Nature Communications, 7:11500. DOI: 10.1038/ncomms11500 

27.        Han, J.*, Hu, S.X., Cartwright, P., Zhao, F.C., Ou, Q., Kubota, S., Wang, X., Yang, X., 2016. The earliest pelagic jellyfish with rhopalia from Cambrian Chengjiang Lagerst?tte. Palaeogeography, Palaeoclimatology, Palaeoecology, 449(2016): 166-173. DOI: 10.1016/j.palaeo.2016.02.025 

28.        杨爱华*, 朱茂炎, 张俊明, 赵方臣, 吕苗. 2015. 扬子板块埃迪卡拉系（震旦系）陡山沱组层序地层划分与对比. 古地理学报,17(1):1-20. 

29.        Yin, Z.J., Zhu, M.Y.*, Davidson E.H., Bottjer D.J., Zhao, F.C., Tafforeau P., 2015. Sponge grade body fossil with cellular resolution dating 60 Myr before the Cambrian. PNAS, 112(12):E1453-E1460. doi:10.1073/pnas.1414577112. 

30.        Zhao, F.C.*, Smith, M. R., Yin, Z.J., Zeng, H., Hu, S.X., Li, G.X., Zhu, M.Y., 2015. First report of Wiwaxia from the Cambrian Chengjiang Lagerst?tte. Geological Magazine, 152(2): 378-382. 

31.        Zeng, H., Zhao, F.C.*, Yin, Z.J., Li, G.X., Zhu, M.Y., 2014. A Chengjiang-type fossil assemblage from the Hongjingshao Formation (Cambrian Stage 3) at Chenggong, Kunming, Yunnan, Chinese Science Bulletin, 59(25): 3169–3175. (封面文章)　 

32.        Zhao, F.C.*, Hu, S.X., Zeng, H., Zhu, M.Y., 2014. A new helmetiid arthropod from the early Cambrian Chengjiang Lagerst?tte, Southwest China. Journal of Paleontology, 88(2): 367–370. 

33.        Zhao, F.C.*, Caron, J.-B., Bottjer, D.J., Hu, S.X., Yin, Z.J., Zhu, M.Y., 2014. Diversity and species abundance patterns of the early Cambrian (Series 2, Stage 3) Chengjiang Biota from China. Paleobiology, 40(1): 50-69. 

34.        Huang, D.Y.*, Chen, J.Y., Zhu, M.Y., Zhao, F.C., 2014. The burrow dwelling behavior and locomotion of palaeoscolecidian worms: New fossil evidence from the Cambrian Chengjiang fauna. Palaeogeography, Palaeoclimatology, Palaeoecology, 398: 154-164. 

35.        胡世学, 朱茂炎, 罗惠麟 Michael Steiner, 赵方臣, 李国祥, 刘琦. 2013. 关山生物群——寒武纪早期海洋生物多样性的再现. 云南科技出版社 (专著) 

36.        Zhao, F.C.*, Bottjer, D.J., Hu, S.X., Yin, Z.J., Zhu, M.Y.,* 2013. Complexity and diversity of eyes in Early Cambrian ecosystems. Scientific Reports 3, 2751; DOI:10.1038/srep02751. 

37.        Zhu, M.Y.*, Lu, M., Zhang, J.M., Zhao, F.C., Li, G.X., Zhao, X., Zhao, M.J., 2013. Carbon isotope chemostratigraphy and sedimentary facies evolution of the Ediacaran Doushantuo Formation in western Hubei, South China. Precambrian Research, 225: 7-28. 

38.        Lu, M., Zhu, M.Y.*, Zhang, J.M., Shields, G.A., Li, G.X., Zhao, F.C., Zhao, X., Zhao, M.J., 2013. The DOUNCE event at the top of the Ediacaran Doushantuo Formation of South China: Wide stratigraphic occurrence and non-diagenetic origin. Precambrian Research，225: 86-109. 

39.        Guo, Q.J.*, Strauss, H., Zhu, M.Y., Zhang, J.M., Yang, X.L., Lu, M., Zhao, F.C., 2013. High resolution organic carbon isotope stratigraphy from a slope to basinal setting on the Yangtze Platform, South China: Implications for the Ediacaran–Cambrian transition. Precambrian Research, 225: 209-217. 

40.        Zhao, F.C.*, Hu, S.X., Caron, J.-B., Zhu, M.Y., Yin, Z.J., Lu, M., 2012. Spatial variation in the diversity and composition of the Lower Cambrian (Series 2, Stage 3) Chengjiang Biota, Southwest China. Palaeogeography, Palaeoclimatology, Palaeoecology, 346-347: 54-65. 

41.        赵方臣*, 朱茂炎, 胡世学, 2012. 不同生活方式的物种对快速沉积埋藏事件的反应—来自寒武纪早期澄江生物群中的证据. 古生物学报, 51(3): 265-280. 

42.        Wang, B.*, Zhao, F.C., Zhang, H.C., Fang, Y., Zheng, D.R., 2012. Widespread pyritization of insects from the Early Cretaceous Jehol biota. Palaios, 27: 707-711. 

43.        Lu, M., Zhu, M. Y.*, Zhao, F.C., 2012. Revisit of the Tianjiayuanzi section: the stratotype section of Ediacaran Doushantuo Formation, Yangtze Gorges, South China. Bulletin of Geosciences, 87(1): 183-194. 

44.        Zhu, M.Y.*, Yang, A.H., Zhao, F.C., Zhang, J.M., Zhao, Y.L., Tai, T.S., 2012. Cryogenian and Cambrian successions in the slope facies of the Yangtze Platform at Wuhe, Jianhe County, Guizhou. In: Zhao, Y., Zhu, M., Peng, J., et al. (eds.), Cryogenian-Ediacaran to Cambrian Stratigraphy and Paleontology of Guizhou, China. Journal of Guizhou University, Natural Sciences. 29 (Sup.1): 98-107. 

45.        Weber, B.*, Hu, S.X., Steiner, M., Zhao, F.C., 2012. A diverse and significant ichnofauna from the Lower Cambrian Wulongqing Formation near Kunming (Yunnan Province, South China). Bulletin of Geosciences, 87(1): 71–92. 

46.        Hu, S.X.*, Steiner, M., Zhu, M.Y., Luo, H.L., Keupp, H., Zhao, F.C., Liu, Q., 2012. A new priapulid assemblage from the early Cambrian Guanshan fossil Lagerst?tte of SW China. Bulletin of Geosciences, 87(1): 93-106. 

47.        Zhao, F.C., Zhu, M.Y.*, Hu, S.X. 2010. Community structure and composition of the Cambrian Chengjiang biota. SCIENCE CHINA Earth Sciences, 53: 1784–1799. [赵方臣，朱茂炎*，胡世学，2010. 云南寒武纪澄江动物群古群落分析。中国科学：地球科学，40（9）：1135-1153] 

48.        Hu, S.X.*, Zhu, M.Y., Steiner, M., Luo, H.L., Zhao, F.C., Liu, Q. 2010. Biodiversity and taphonomy of the Early Cambrian Guanshan biota, eastern Yunnan. SCIENCE CHINA Earth Sciences, 53: 1765–1773. [胡世学*，朱茂炎，Steiner, M., 罗惠麟，赵方臣，刘琪 2010. 关山生物群构成及埋藏学研究进展。中国科学：地球科学，40（9）：1115-1124 ] 

49.        王向东，王玥，张以春，赵方臣，盛青怡. 2010. 古生态学、古气候学、古生物地理学. 古生物学学科发展报告，中国科学技术协会主编，中国科学技术出版社出版，63-69. 

50.        Zhao, F.C., Caron, J.B., Hu, S.X., Zhu, M.Y.* 2009. Quantitative analysis of Taphofacies and Paleocommunities in the early Cambrian Chengjiang Lagerst?tte. Palaios, 24: 826-839. (Cover article) 

51.        赵方臣, 朱茂炎*. 2007.云南早寒武世澄江化石库中两种埋藏相的化石定量分析. 古生物学报, 46(1): 75-86. 

52.        Zhu, M.Y., Li, G.X., Hu, S.X., Zhao, F.C. 2005. Ediacaran-Cambrian boundary stratigraphy at Meishucun, Jinning County, and fossil quarries of the Early Cambrian Chengjiang Biota Haikou, Kunming city, Yunnan, China. In: Peng S C, Babcock L E, Zhu M Y (eds). Cambrian System of China and Korea. University of Science and Technology of China Press. 13-29. 
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