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Yuan Xunlai
Research direction:
Postal Code:
39 East Beijing Road, Nanjing 210008, China


Personal information 

  Sex: Male 

  Birthday: August, 1, 1965 

  Address:   Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences. No.39 East Beijing Road, Nanjing 210008, CHINA 

  Phone: 025-83282217 

  Fax: 025-83357026 

  E-mail: xlyuan@nigpas.ac.cn 


  Peking University       Geology        M. Sc., 1990 

  Peking University       Geology        B. Sc., 1987 

Current Position and Professional Experience 

  2000 – present   Director, Department of Micropaleontology, Nanjing Institute of Geology and Palaeontology, CAS 

  2001 – present  Professor               Nanjing Institute of Geology and Palaeontology, CAS 

  1998 – 2001   Associate Professor      Nanjing Institute of Geology and Palaeontology, CAS 

  1993 – 1998   Research Associate      Nanjing Institute of Geology and Palaeontology, CAS 

  1990 – 1993   Research Assistant       Nanjing Institute of Geology and Palaeontology, CAS 

  1999 – 2000  Academic Visitor     Organismic & Evol. Biology,     Harvard University 

  1995 – 1996  Academic Visitor     Department of Geology,    University Of Montreal 

Areas of Interest 

  Fields: the origin and early evolution of multicellular organisms (多细胞生物起源和早期演化) 

Awards and Honors 

  Jiangsu Science and Technology Progress Award (2013) 

  Ten Pieces of News in Science and Technology of 2008 in China 

  Special Government Allowance (2007) 

  National Science Fund for Distinguished Young Scholars (2006) 

  Yin Zanxun Stratigraphy and Paleontology Award (2005) 

  Science Chinese, Person of the Year 2005 

  Ten Pieces of News in Science and Technology of 2005 in China 

  New Century National Talent Project (2004) 

  China Book Award (2004) 

  Best Paper Award, Journal of Paleontology (2003) 

Selected Synergistic Activities 

  (a) Professional Services: International Subcommission on Ediacaran Stratigraphy (Voting member, 2012-present), Palaeontological Society of China (Standing Committee, 2005-present), Micropalaeontological Society of China (Chair, 2004-2012), Fossil Algae Society of China (Chair, 2005-2012). 

  (b) Chief and Associate Editorship: Science Bulletin (Earth Science Associate editor); Acta Micropalaeontologica Sinica; Acta Palaeontologica Sinica (Associate editor); Journal of Stratigraphy; Evolution of Llife (Chief editor). 

Collaborators and Other Affiliations 

  (a) Collaborators in past five years: Xiao, S. (Virginia Tech); Bao H. (Louisiana State University); A. J. Kaufman (Univ. Maryland); J. D. Schiffbauer (Univ. Missouri); T. Lyons (UC Riverside); Hua H., (Northwest University); Yin L., Zhou C., & Chen Z. (Chinese Academy of Sci.). 

  (b) Graduate and Postdoctoral Advisors: H.J. Hofmann (McGill University, deceased); Y. Zhang (Peking University, deceased). 

  (c) Thesis Advisor and Postgraduate-Scholar Sponsor: 

  Current graduate students: Wan Bing (PhD), Chen Lei (PhD), Dai Zhen (PhD), Pang Ke (PhD), Ou Zhiji ((MSc), Cai Yaoping (post-doc), Wang Yuliang ((visiting scientist). 

  Past students and post-doctoral fellows: Meng Fanwei (MSc and PhD, now at Nanjing Institute of Geology and Palaeontology, CAS), Hu Jie (MSc and PhD, now at China National Petroleum Corporation), Kong Fanfan (PhD, now at China University of Mining and Technology), Wang Dan (MSc and PhD, now at Nanjing Institute of Geology and Palaeontology, CAS), Peng Yongbo (MSc, now at Indiana University), Yan Xianqing (MSc, now at Geological Survey of Jiangsu Province), Xiang Ke (MSc, now at Hunan Institute of Geological Survey), Song Weiming (MSc, now at East China Mineral Exploration and Development Bureau), Qian Maiping (post-doc; now at Geological Survey of Jiangsu Province), Hua Hong (post-doc; now at Northwest University). 


  1. Zhang Yun and Yuan Xunlai, 1992: New data on multicellular thallophyte and fragments of cellular tissues from Late    Proterozoic phosphate rocks, South China. Lethaia, 25:1-18. Pl.10, Appendix 1. 

  2. 袁训来,王启飞,张昀,1993:贵州瓮安磷矿晚前寒武纪陡山沱期的藻类化石群。微体古生物学报 104):409-420。图3,图版5 

  3. 袁训来,李军,陈孟莪,1995:晚前寒武纪后生植物的发展及其化石证据。古生物学报 341):90-102。图3,图版3 

  4. 张昀,袁训来,1995:元古宙末多细胞红藻有性生殖结构的发现。中国科学(B辑)257):749-754。图3,图版2(Zhang Yun and Yuan Xunlai, 1996: Sexual reproductive structures of Latest Proterozoic multicellular rhodophytes (red algae) from South China. Science in China (Series C) 39(1): 28-39.) 

  5. Yuan Xunlai and H.J. Hofmann, 1998: New micropaleontological data from Neoproterozoic Sinian Doushantuo phosphorite rocks,   Wengan, Guizhou Province, southwestern China. Pl. 14, Appendix 1. Alcheringa 22(3-4): 189-222. 

  6. Shuhai Xiao, A.H. Knoll and Xunlai Yuan, 1998: Morphological reconstruction of Miaohephyton bifurcatum, a possible brown alga from the Neoproterozoic Doushantuo Formation, South China. Journal of Paleontology 72(6):1072-1086. 

  7. Zhang Y., Yuan X. & Yin L. 1998: Interpreting Late Precambrian Microfossils. Science 282(5395):1783. 

  8. 袁训来,1999:新元古代陡山沱期瓮安生物群研究简况。《微体古生物学报》163):281-286 

  9. Yuan Xunlai, Li Jun &Cao Ruiji,1999: A diverse metaphyte assemblage from the Neoproterozoic black shales of South China. Lethaia 32:143-155 

  10. Xiao Shuhai, Yuan Xunlai,A.H. Knoll, 2000: Eumetazoan Fossils in Terminal Proterozoic Phosphorites? PNAS (美国科学院院报)9725):13684-13689 

  11. Yuan Xunlai, Xiao Shuhai, Li Jun, Yin Leiming, and Cao Ruiji2001: Pyritized chuarids with excystment structures from the late Neoproterozoic Lantian Formation in Anhui, South China. Precambrian Research 107(3-4): 253-263. 

  12. 曹瑞骥,袁训来,肖书海,2001:论锥叠层石群(Conophyton)的形态发生──对苏北新元古代九顶山组一个似锥叠层石的剖析。古生物学报403):318-329 

  13. Shuhai Xiao, Xunlai Yuan, Michael Steiner, and A. H. Knoll, 2002: Macroscopic carbonaceous compressions in a terminal Proterozoic shale: A systematic reassessment of the Miaohe Biota, South China. Journal of Paleontology, 76(2): 347-376.  

  14. 袁训来,肖书海,尹磊明,A.H. Knoll,周传明,穆西南:陡山沱期生物群:早期动物辐射前夕的生命。(专著)171页,中国科技大学出版社,合肥。2002.7 

  15. Zhou Chuanming, Yuan Xunlai, Xiao Shuhai, 2002: Phosphatized biotas from the Neoproterozoic Doushantuo Formation on the Yangtze Platform. Chinese Science Bulletin, 47(22): 1918-1924. 

  16. 胡杰,陈哲,薛耀松,王金权,王金龙,袁训来,2002:皖南早寒武世荷塘组海绵骨针化石。微体古生物学报,191):53-62 

  17. Yuan Xunlai, Shuhai Xiao, Ronald L. Parsley, Chuanming Zhou, Zhe Chen, Jie Hu, 2002: Towering sponges in an Early Cambrian Lagersta¨tte: Disparity between nonbilaterian and bilaterian epifaunal tierers at the Neoproterozoic-Cambrian transition. Geology, 30(4): 363-366. 

  18. Xiao, S., J. Hu, X. Yuan, R. L. Parsley, and R. Cao, 2005: Articulated sponges from the Lower Cambrian Hetang Formation in southern Anhui, South China: Their age and implications for the early evolution of sponges. Palaeogeography, Palaeoclimatology, Palaeoecology, 220:89-117. 

  19. Xiao, S., Huiming Bao, Haifeng Wang, A J Kaufman, Chuanming Zhou, Guoxiang Li, Xunlai Yuan, Hongfei Ling, 2004, The Neoproterozoic Quruqtagh Group in eastern Chinese Tianshan: evidence for a post-Marinoan glaciation. Precambrian Research, 130, 1-26. 

  20. 尹磊明,边立曾,袁训来,2003:山西中元古代汝阳群分枝管状藻体和显微环螺旋加厚管体的发现。中国科学(D),338):769-774 

  21. Xiao, S., A. H. Knoll, X. Yuan, and C. M. Pueschel, 2004, Phosphatized multicellular algae in the Neoproterozoic Doushantuo Formation, China, and the early evolution of florideophyte red algae. American Journal of Botany, 91:214-227. 

  22. 孟凡巍袁训来 周传明,2003:新元古代大塘坡组黑色页岩中的甲藻甾烷及其生物学意义。微体古生物学报,194):97-102 

  23. Cao Rui-ji, Yuan Xun-lai and Mu Xi-nan,2003: Brief History and Current Status of Stromatolite Study in China. 微体古生物学报,194):5-14 

  24. Yin Leiming, Yuan Xunlai, 2003: A review of eukaryotic fossils in the late Mesoproterozoic Ruyang Group in Shanxi, Chin. 微体古生物学报,194):39-46 

  25. Mu Xi-nan, Yuan Xun-lai, 2003: Introduction of Fossil algae and Stromatolites. 微体古生物学报,194):1-4 

  26. Bao, Huiming; Thiemens, Mark H.; Loope, David B.; Yuan, Xun-Lai, 2003: Sulfate oxygen-17 anomaly in an Oligocene ash bed in mid-North America: Was it the dry fogs? Geophys. Res. Lett.Vol. 30, No. 16, 1843 

  27. Chuanming Zhou, R Tucker, Shuhai Xiao, Zhanxiong Peng, Xunlai Yuan, Zhe Chen, 2004, New constraints on the ages of Neoproterozoic glaciations in south China. Geology, 32(5): 437-440. 

  28. 钱迈平,袁训来等,2002:苏、皖北部新元古代微生物化石。微体古生物学报,194):363-381 

  29. Cao Rui-ji and Yuan Xun-lai, 2003: Pre-Sinian biostratigraphy of China. In: Zhang et al. eds. Biostratigraph of China. 1-30. Science Press, Beijing. 

  30. Yin Lei-ming and Yuan Xun-lai, 2003: Sinian biostratigraphy of China. In: Zhang et al. eds. Biostratigraphy of China. 31-54. Science Press, Beijing. 

  31. Chen Zhe, Hu Jie, Zhou Chuanming, Xiao Shuhai, Yuan Xunlai, 2004, Sponge assemblage from early Cambrian Hetang Formation, southern Anhui. Chinese Science Bulletin, 49(15): 1625-1628. 

  32. Yin Leiming, Bian Lizeng, Yuan Xunlai, 2004, Discovery of branched tubular algae and microscopic tubes with annular-helical thickening from the Mesoproterozoic Ruyang Group of Shanxi, North China. Science in China (Ser. D), 47(10): 880-885. 

  33. Hua Hong, Chen Zhe, Yuan Xunlai, Xiao Shuhai, Zhang Luyi, 2005. Skeletogenesis and asexual reproduction in the earliest biomineralizing animal Cloudina. Geology, 33(4): 277-280. 

  34. 周传明,袁训来,肖书海,陈哲,薛耀松,2004:湖北保康陡山沱磷酸盐化微体化石组合。微体古生物学报,214):349-366 

  35. Jinxian Yao, Shuhai Xiao, Leiming Yin, Guoxiang Li and Xunlai Yuan,2005:Basal Cambrian microfossils from the Yurtus and Xishanblaq formations (Tarim, north-west China): Systematic revision and biostratigraphic correlation of Micrhystridium-like acritarchs from China. Paleontology, 48(4): 687-708 

  36. Shuhai Xiao, Jie Hu, Xunlai Yuan, Ronald L. Parsley, Ruiji Cao2005Articulated sponges from the Lower Cambrian Hetang Formation in southern Anhui, South China: their age and implications for the early evolution of sponges. Palaeogeography, Palaeoclimatology, Palaeoecology 220: 89-117 

  37. 周忠和,骆永明,董云社,袁训来,高星,2005:浅谈我国地球环境与生命过程研究中的一些科学问题。科学通报,502):195-200 

  38. Shuhai XiaoBing ShenChuanming Zhou Guwei Xie Xunlai Yuan2005: A uniquely preserved Ediacaranfossil with directevidence for a quilted bodyplan. PNAS, 102(29): 10227-10232. 

  39. Xunlai Yuan, Shuhai Xiao, T.N., Taylor, 2005: Lichen-like symbiosis 600 million years ago. Science, 308: 1017-1020. 

  40. Meng Fanwei, Zhou Chuanming, Chen Zhilin, Yuan Xunlai, 2005: The oldest known dinoflagellates: morphological and molecular evidence from Mesoproterozoic rocks at Yongji, Shanxi Province. Chinese Science Bulletin, 50(12): 1230-1234. 

  41. Yin Leiming, Yuan Xunlai , Meng Fanwei and Fu Jie, 2005: Protists of the upper Mesoproterozoic Ruyang Group in Shanxi Province, China. Precambrian Research, 141: 29-66. 

  42. 孟凡巍,周传明,燕夔,袁训来,尹磊明,2006:通过C27/C29甾烷和有机碳同位素来判断早古生代和前寒武纪的烃源岩的生物来源。微体古生物学报,231):51-56 

  43. James W. Hagadorn1, Shuhai Xiao, Philip C. J. Donoghue, Stefan Bengtson, Neil J. Gostling, MariaPawlowska, Elizabeth C. Raff5, Rudolf A. Raff, F. Rudolf Turner, Yin Chongyu, Chuanming Zhou, XunlaiYuan, Matthew B. McFeely, Marco Stampanoni, Kenneth H. Nealson2006Cellular and Subcellular Structure of Neoproterozoic Animal Embryos. Science 314291-294 

  44. Shuhai Xiao, James W. Hagadorn, Chauming Zhou, Xunlai Yuan, 2007: Rare helical spheroidal fossils from the Neoproterozoic Doushantuo lagerstätte: Ediacaran animal embryos come of age? Geology 35: 115-118. 

  45. 袁训来,肖书海,周传明,2006:新元古代陡山沱期真核生物的辐射。戎嘉余(主编):《生物的起源、辐射与多样性演变-华夏化石记录的启示》。北京:科学出版社。13-27 

  46. 曹瑞骥,袁训来,2006:《叠层石》。合肥:中国科技大学出版社。383页。 

  47. Leiming Yin, Maoyan Zhu, Andrew H. Knoll, Xunlai Yuan, Junming Zhang & Jie Hu2007Doushantuo embryos preserved inside diapause eggcysts. Nature 445, 198201 

  48. Shuhai Xiao, Chuanming Zhou,Xunlai Yuan 2007: Undressing and redressing Ediacaran embryos Nature 445, 198201. 

  49. ZHOU CHUANMING, XIE GUWEI, KATHLEEN MCFADDEN, XIAO SHUHAIand YUAN XUNLAI 2007: The diversification and extinction of Doushantuo-Pertatatakaacritarchs in South China: causes and biostratigraphic significance. GEOLOGICAL JOURNAL42: 229262. 

  50. HUA HONG, CHEN ZHE and YUAN XUNLAI, 2007The advent of mineralized skeletons in Neoproterozoic Metazoanew fossil evidence from the Gaojiashan Fauna. GEOLOGICAL JOURNAL42: 263279. 

  51. Lin Dong, Shuhai Xiao, Bing Shen, Xunlai Yuan, Xianqin Yan, Yongbo Peng,2008Restudy of the worm-like carbonaceous compression fossils Protoarenicola, Pararenicola, and Sinosabellidites from early Neoproterozoic successions in North China, Palaeogeography, Palaeoclimatology, Palaeoecology 258 138161 

  52. Yin Leiming, Yuan Xunlai, 2007: Radiation of Meso-Neoproterozoic and early Cambrian protistsinferred from the microfossil record of China. Palaeogeography, Palaeoclimatology, Palaeoecology, 254: 350361 

  53. James D. S., Leiming Y, Robert J. B., A. J. Kaufman, Fanwei M., Jie H., Bing S. Xunlai Y., Huiming B.,and Shuhai X., 2007: Ultrastructural and Geochemical Characterization ofArcheanPaleoproterozoic Graphite Particles:Implications for Recognizing Traces of Life in HighlyMetamorphosed Rocks. Astrobiology 7(4): 684-704. 

  54. 严贤勤, 孟凡巍,袁训来, 2006:徐淮地区新元古代九顶山组燧石结核的地球化学特征。微体古生物学报, 233);295-302 

  55. 袁训来,2007:话说六亿年前的动物胚胎化石。生物进化,116-19. 

  56. 彭永波,王丹,袁训来,2007;长城系串岭沟组球形疑源类的超微结构。微体古生物学报, 242);194-204 

  57. 孟凡巍、袁训来等,2007:同步辐射X射线荧光微探针技术测定太古宇条带状硅铁建造中硅质条待鸡包裹体的微量元素。岩石学报,(092079-2084。(SCI 

  58. Kathleen A. M., Jing Huang, Xuelei Chu, Ganqing Jiang,Alan J.Kaufman, Chuanming Zhou, Xunlai Yuan and Shuhai Xiao, 2008: Pulsed oxidation and biological evolution in the Ediacaran Doushantuo Formation. PNAS 105(9): 3197-3202. 

  59. 尹磊明,周传明,袁训来,2008:湖北宜昌埃迪卡拉系陡山沱组天柱山乱囊胞——Tianzhushania的新认识。古生物学报,472):129-140.  

  60.    Yongbo Peng, Huiming Bao, Xunlai Yuan,2009: New morphological observations for Paleoproterozoic acritarchsfrom the Chuanlinggou Formation, North China. Precambrian Research168223–232. 

  61.    Bei Xu, Shuhai Xiao, Haibo Zou, Yan Chen, Zheng-Xiang Li , Biao Song, Dunyi Liu,Chuanming Zhou, Xunlai Yuan2009SHRIMP zircon U–Pb age constraints on Neoproterozoic Quruqtaghdiamictites in NW China.Precambrian Research168247–258. 

  62.    袁训来,王丹,肖书海,2009:新元古代陡山沱期的动物。古生物学报,48(3):375-389. 

  63.    曹瑞骥,袁训来,2009:中国叠层石研究进展。古生物学报,48(3):314-321. 

  64.    Shen, B., Xiao, S., Zhou, C., Yuan, X., 2009, Yangtziramulus zhangi new genus and species, a carbonate-hosted Ediacaran macrofossil from the Dengying Formation in the Yangtze Gorges area, South China, Journal of Paleontology, 83: 575-587. 

  65.    Bing Shen, Shuhai Xiao, Chuanming Zhou, Alan J. Kaufmand, Xunlai Yuan2010Carbon and sulfur isotope chemostratigraphy of the Neoproterozoic QuanjiGroup of the Chaidam Basin, NW China: Basin stratification in theaftermath of an Ediacaran glaciation postdating the Shuram event?Precambrian Research 177 241–252. 

  66.    Zhou Chuanming, Bao Huiming, Peng Yongbo, Yuan Xunlai, 2010, Timing the deposition of 17O-depleted barite at the aftermath of Nantuo glacial meltdown in South China. Geology, 38: 903-906. 

  67.    华洪, 陈哲, 袁训来, 肖书海, 蔡耀平, 2010, 陕南伊迪卡拉纪末期的瓶状化石——可能最早的有孔虫化石: 中国科学,D, v. 40, no. 9, p. 1105-1114. 

  68.    龚淼, 周传明, 尹磊明, 袁训来, 王春朝, 2010, 元古代刘老碑组Leiosphaeridia asperata膜壁的超微结构及其亲缘关系探讨。中国科学,D, 409):1098-1104. 

  69.    GONG Miao, ZHOU ChuanMing, YIN LeiMing, YUAN XunLai & WANG ChunZhao, 2010: Cell wall ultrastructures of the Proterozoic acritarch Leiosphaeridia asperata and their implications for biological affinity. Science in China Series D, 53 (12): 1750-1755 

  70.    Fanwei Meng, Pei Nia, James D. Schiffbauerc, Xunlai Yuan, Chuanming Zhou,Yigang Wang, Maolong Xia2011: Ediacaran seawater temperature: evidence from inclusions of Sinian halite. Precambrian Research 184 (1-4): 63-69. 

  71.    Xunlai Yuan, Zhe Chen, Shuhai Xiao, Chuanming Zhou, HongHua, 2011: An early Ediacaran assemblage of macroscopic and morphologically differentiated eukaryotes. Nature, 470: 390-393.  

  72.    Yongbo Peng, Huiming Bao, Chuanming Zhou, Xunlai Yuan, 2011: 17O-depleted barite from two Marinoan cap dolostone sections, South China. Earth and Planetary Science Letters 305:  21–31. 

  73.    王鑫, 袁训来, 周传明, 杜开和, 龚淼, 2011: Chuaria的解剖结构及其植物属性。科学通报,569):693 ~ 699 

  74.    Shen, B., S. Xiao, H. Bao, A. J. Kaufman, C. Zhou, and X. Yuan,2011Carbon and sulfur isotope evidence for a strong depth gradient and oceanic oxidation after the Ediacaran Hankalchough glaciation. Geochimica et Cosmochimica Acta, 75: 1357-1373. 

  75.    WANG Xin, YUAN XunLai, ZHOU ChuanMing, DU KaiHe& GONG Miao2011Anatomy and plant affinity of Chuaria. Chinese Sci Bull, 56: 1256−1261 

  76.    Hua, Hong, Chen, Zhe; Yuan, XL, et al.,2010: The earliest Foraminifera from southern Shaanxi, China. SCIENCE CHINA-EARTH SCIENCES, 53 (12): 1756-1764. 


  77.    Lei-Ming Yin, Dan Wang, Xun-Lai Yuan, Chuan-Ming Zhou, 2011: Diverse small spinose acritarchs from the Ediacaran Doushantuo Formation,South China. Palaeoworld 20: 279–289. 

  78.    KONG FanFan, YUAN XunLai & ZHOU ChuanMing, 2011: Paleoproterozoic glaciation: Evidence from carbon isotope record ofthe Hutuo Group, WutaiMountain area of Shanxi Province, China. Chinese Sci Bull, 56: 2922−2930. 

  79.    Shuhai Xiao, Andrew H. Knoll, James D. Schiffbauer, Chuanming Zhou, Xunlai Yuan, 2012: Comment on “Fossilized Nuclei andGermination Structures Identify Ediacaran‘Animal Embryos’ as Encysting Protists” Science 3351169a-c. 

  80.    ZHOU ChuanMing, JIANG ShaoYong, XIAO ShuHai, CHEN Zhe & YUAN XunLai, 2012: Rare earth elements and carbon isotope geochemistry of theDoushantuo Formation in South China: Implication formiddle Ediacaran shallow marine redox conditions. Chin Sci Bull, doi: 10.1007/s11434-012-5082-6 

  81.    Wei Wang, Chuanming Zhou, Xunlai Yuan, Zhe Chen, Shuhai Xiao, 2012:A pronounced negative δ13C excursion in an Ediacaran succession of western YangtzePlatform: A possible equivalent to the Shuram event and its implication forchemostratigraphic correlation in South China. Gondwana Research 22: 1091–1101. 

  82.    Zhe Chen, Chuanming Zhou, Mike Meyer, Ke Xiang, James D. Schiffbauerd, Xunlai Yuan,Shuhai Xiao2013Trace fossil evidence for Ediacaran bilaterian animals with complex behaviors. Precambrian Research 224 (2013) 690– 701 

  83.    袁训来, 陈哲, 肖书海, 万斌, 关成国, 王伟, 周传明, 华洪,2012:蓝田生物群: 一个认识多细胞生物起源和早期演化的新窗。科学通报5734):3219 ~ 3227 

  84.    Yuan, X., Chen, Z., Xiao, S., Wan, B., Guan, C., Wang, W., Zhou, C.,and Hua, H., 2013, The Lantian biota: A new window onto the originand early evolution of multicellular organisms: Chinese ScienceBulletin, 58: 701-707.  

  85.    代珍,严贤勤,袁训来,尹磊明,2012:徐淮地区元古宙九顶山组燧石中微体生物化石新资料。古生物学报,51(1): 26-55 

  86.    Yongbo Peng, Huiming Bao, Chuanming Zhou, Xunlai Yuan, Taiyi Luo, 2013, Oxygen isotope composition of meltwater of a Neoproterozoic glaciation in South China. Geology, 41:367-370. 

  87.    董琳,宋伟明,肖书海,袁训来,陈哲,周传明,2012,皖南地区埃迪卡拉系皮园村组微体和宏体化石兼论埃迪卡拉系寒武系界线。地层学杂志,36(3): 600-610 

  88.    Shuhai Xiao, Mary Droser, James G. Gehling, Ian V. Hughes, Bin Wan, Zhe Chen, and Xunlai Yuan2013, Affirming life aquatic for the Ediacara biota in China and Australia: Geology, v. 41, no. 10, p. 1095-1098. 

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