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Wang Xin
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Postal Code:
39 East Beijing Road, Nanjing, 210008, China



1986-1990  Department of Geology, Peking University, BS

1990-1993  Institute of Botany, CAS, MS (Dr. Ye Chen and Dr. Shuying Duan)

1998-2004  University of Florida, PhD (Dr. David Dilcher (USA academy member))

2004-2006 Nanjing Institute of Geology and Palaeontology, CAS, Postdoctor (Dr. Zhiyan Zhou (Chinese academy member))



Institute of Botany, Chinese Academy of Sciences            1993- 1997    Researcher Assistant.

Hainan Minfu Investment Group, Hainan, PRC                   1995-1997     Project Manager

Unilation Communications Company, Beijing, PRC            1997-1998     Manager

Paleobotany Lab, FLMNH, Florida, USA                             1998-2000     Researcher Assistant

Geology Department, UF, USA                                              2000-2001     Collection Curator

Geology Department, UF, USA                                              2001-2002     Teaching Assistant

Disorder Education, UF, USA                                                2001-2001     Culture Consultant

Geology Department, UF, USA                                             2002-2004     Lab Technician

Nanjing Institute of Geology and Palaeontology               2004-2006     Postdoctor

Nanjing Institute of Geology and Palaeontology               2006-2011     Associate Research Professor

Nanjing Institute of Geology and Palaeontology               2011-             Research Professor



Origin, radiation and development of angiosperms; Fossil cytoplasm of plants and its ultrastructures.


  1990  Award for Excellent Graduate, Peking University

  1991  Award for Excellent Graduate, Graduate School, Chinese Academy of Sciences

  1999  Award of Outstanding Academic Achievement by an International Student, University of Florida

  2000  Award of Outstanding Academic Achievement by an International Student, University of Florida

  2002  Award of Outstanding Academic Achievement by an International Student, University of Florida

  2002  Presidential Recognition of Outstanding Achievement and Contribution to University of Florida

  2010  Included in Marquis Who’sWho in America

  2011  Included in the 333 Program of Jiangsu Province


Peer-reviewed Papers

1995, Wang X. Study on the middle Jurassic flora of Tongchuan, Shaanxi Province. Chinese Journal of Botany, 7(1):81-88.

1995, Duan SY, Cui JZ, Wang X, Xiong BQ, Wang YQ. Fossil woods from the early Cretaceous of western Liaoning, China, In Wu SM. Wood Anatomy Research, International Symposium on Tree Anatomy and Wood Formation, Tianjin, China. Tianjin International Academic Publishers, 166-171.

1997, Duan S, Wang X. A peculiar fossil leaf Beipiaophyllum Duan & Wang gen. nov., Chenia, 3-4:125-131

1997, Wang X, Duan S, Cui J. Several species of Schizolepis and their significance on the evolution of conifersTaiwania, 42(2):73-85.

1997, Xiong B, Wang Y, Duan S, Cui J, Wang X. Rare earth elements in the woodstone.  Journal of the Chinese Rare Earth Society, Special Issue, 15:428-431

2001, Jarzen DM, Wang X. Book review of “Fossil spores and pollen of China”, AASP News, 21(3):8-9.

2004, Wang X. Plant cytoplasm preserved by lightning, Tissue & Cell, 36(5):351-360.

2006, Wang X., A chemical signal possibly related to physiology in fossil cells detected by energy dispersive X-ray microanalysis, Tissue & Cell, 38(1):43-51.

2006Wang X. Li N, Cui J. Jurastrobus chenii gen. et sp. nov., a cycadalean pollen cone connected with vegetative parts from Inner Mongolia, China, Progress of Natural Sciences, 16(13):213-221.

2007, Wang X. High temperature as a mechanism for plant cytoplasm preservation, Acta Geologica Sinica, 81(2):183-193.

2007, Wang X., Cui, J., The first observation on plant cell fossils in China. Acta Geologica Sinica, 81(1):16-22.

2007, Wang X. Duan S, Geng B, Cui J, Yang Y. Schmeissneria: A missing link to angiosperms? BMC Evolutionary Biology, 7:14.

2007, Wang X. Vivid fossils, Evolution of Life, 1:20-22.

2007, Wang X, Liu WZ, Cui JZ, Du KH. Palaeontological evidence for membrane fusion between a unit membrane and a half unit membrane, Molecular Membrane Biology, 24(5-6):496-506.

2007, Wang X, Duan S, Geng B, Cui J, Yang Y. Is Jurassic Schmeissneria an angiosperm? Acta Palaeontologica Sinica, 46(4):486-490.

2008, Wang X. Mesofossils with platanaceous affinity from the Dakota Formation (Cretaceous) in Kansas, USA. Palaeoworld, 17, 246-252.

2008, Wang X, Yu J, Fang X. An AFM observation on fossil cytoplasm. Acta Geologica Sinica, English version, 82, 1141-1145.

2008, Wang X. Dilcher DL, Lott T, Li Y. Parapodocarpus gen. nov. and its implications for interpreting the ovulate organ in the Podocarpaceae. Geophytology, 37:1-8.

2009, Wang X, Li N, Wang Y, Zheng S. The discovery of whole-plant fossil cycad from the Upper Triassic in western Liaoning and its significance. Chinese Science Bulletin 54: 3116-3119.

2009, Wang X, Li N, Wang Y, Zheng S. The discovery of whole-plant fossil cycad from the Upper Triassic in western Liaoning and its significance. Kexue Tongbao, 54:1937-1939.

2009, Wang X, Zheng S. The earliest normal flower from Liaoning Province, China. Journal of Integrative Plant Biology 51: 800-811.

2009, Jin J, Kodrul TM, Liao W, Wang X. A new species of Craigia from the Eocene Changchang Formation of Hainan Island, China. Review of Palaeobotany and Palynology 155:80-82.

2010. Wang X. Axial nature of cupule-bearing organ in Caytoniales. Journal of Systematics and Evolution 48(3):207-214.

2010. Wang X. Schmeissneria: An angiosperm from the Early Jurassic. Journal of Systematics and Evolution 48(5):226-235.

2010. Wang X, Krings M, Taylor TN. A thalloid organism with possible lichen affinity from the Jurassic of northeastern China. Review of Palaeobotany and Palynology 162:567--574.

2010. Wang X., and Wang S. Xingxueanthus: an enigmatic Jurassic seed plant and its implications for the origin of angiospermy. Acta Geologica Sinica 84(1):47-55.

2010. Wang X., Zhang Y, Du K, Fang X. Atomic force microscope observation on ultrastructures in plant cells. Journal of Nanoscience and Nanotechnology 10(10):6624-6628.

2010. Wang X, Zheng SL. Whole fossil plants of Ephedra and their implications on the morphology, ecology and evolution of Ephedraceae (Gnetales). Chinese Science Bulletin 55(15):1511-1519.

2010. Wang X, Zheng SL, Jin JH. Structure and relationships of Problematospermum, an enigmatic seed from the Jurassic of China. International Journal of Plant Sciences 171(4):447-456.

2010. Wang Y, Liu Z, Wang X, Shih C, Zhao Y, Engel MS, Ren D. Ancient pinnate leaf mimesis among lacewings. Proceedings of the National Academy of Sciences 107(37):16212-16215.

2010. Wang X, Du K, Yi T, Jin J. Cytoplasmic remains in an Eocene fossil stem. IAWA Journal 31(3):363-367.

2010. Liu WZ, Zhou YF, Wang X, Jiao ZJ. Programmed cell death during pigment gland formation in Gossypium hirsutum leaves. Plant Biology 12(6):895-902.

2010. Zheng S, Wang X. An undercover angiosperm from the Jurassic in China. Acta Geologica Sinica 84(4):895-902.

2011. Wang X, Liu W, Du K. Palaeontological evidence of membrane relationship in step-by-step membrane fusion. Molecular Membrane Biology, 28:115-122.

2011. Wang X, Yuan X, Zhou C, Du K, Gong M. Anatomy and plant affinity of Chuaria. Chinese Science Bulletin 56:1256-1261

2011. Wang, X., Han, G. The earliest ascidiate carpel and its implications for angiosperm evolution. Acta Geologica Sinica 85:998-1002.

2012. Wang X, Zheng X-T. Reconsiderations on two characters of early angiosperm Archaefructus. Palaeoworld 21: 193-201

2012. Tanner LH, Wang X, Morabito AC. Fossil charcoal from the Middle Jurassic of the Ordos Basin, China and its paleoatmospheric implications. Geoscience Frontiers 3: 493-502

2013. Han G, Fu X, Liu Z-J, Wang X. A new angiosperm genus from the Lower Cretaceous Yixian Formation, Western Liaoning, China. Acta Geologica Sinica 87, 916-925.

2013. Li N, Deng H-X, Zhang W-H, Du K-H, Wang X. The impact of high temperature on living Plant cytoplasm. Acta Palaeontologica Sinica 52:484-491.

2013. Wang X, Luo B. Mechanical pressure, not genes, makes ovulate parts leaf-like in Cycas. American Journal of Plant Sciences 4(12A):53-57.

2014. Heinrichs J., Wang X., Ignatov M.S. and Krings M. A Jurassic moss from Northeast China with preserved sporophytes. Review of Palaeobotany and Palynology, 204: 50-55.

2014. Wang, X., Liu, W., Du, K., He, X. and Jin, J. Ultrastructure of chloroplasts in fossil Nelumbo from the Eocene of Hainan Island, South China. Plant Systematics and Evolution, 10:2259-2264.

2015. Wang X. The megafossil record of early angiosperms in China since 1930s. Historical Biology 27:398-404

2015. Wang, X. Progress in the study on plant cytoplasm fossils. Science & Technology Review 33(7):108-113.

2015. Wang, X., Liu, Z.-J., Liu, W., Zhang, X., Guo, X., Hu, G., Zhang, S., Wang, Y., Liao, W. Breaking the stasis of current plant systematics. Science & Technology Review. 33(22):97-105.

2015. Wang, X., Liu, Z.-J., Jurassic flowers and the origin of angiosperms. Chinese Journal of Nature 37(6): 435-440.

2016. Han, G., Liu, Z.-J., Liu, X., Mao, L., Jacques, F.M.B., Wang, X. A whole plant herbaceous angiosperm from the Middle Jurassic of China. Acta Geologica Sinica 90(1):19-29.

2016. Liu, Z.-J., Wang, X. A perfect flower from the Jurassic of China. Historical Biology 28(5):707-719.

2016. Liu, Z.-J., Wang, X. An enigmatic Ephedra-like fossil lacking micropylar tube from the Lower Cretaceous Yixian Formation of Liaoning, China. Palaeoworld 25(1):67-75.

2016. Shen, H.-X., Du, K.-H., Wang, X. Plant cytoplasm preservation in a baked root of Abies. Palaeoworld 25: 287-291.

2016. Wang, X. On the interpretation of fossil nuclei. Natural Science 8: 216-219.

2017. Liu, Z.-J., Wang, X. Yuhania: A unique angiosperm from the Middle Jurassic of Inner Mongolia, China. Historical Biology, 29(4):431-441

2017. Zhang, X., Liu, W., Wang, X., How the ovules get enclosed in magnoliaceous carpels? PLOS One 12(4): e0174955.

2017. Han, G., Liu, Z.-J., Wang, X., A Dichocarpum-like angiosperm from the Early Cretaceous of China. Acta Geologica Sinica (English Edition), 91(1):1-8

2017. Wang, X., A biased, misleading review on early angiosperms. Natural Science, 9(12): 399-405.

2017. Miao, Y., Liu, Z. J., Wang, M., and Wang, X., Fossil and living cycads say "No more megasporophylls". Journal of Morphology and Anatomy, 1(2):1000107.

2017. Wang, X., Do life and prime numbers all have genes. Evolution, 9(12): 399-405.

2018. Liu, Z.-J., Huang, D., Cai, C., Wang, X., The core eudicot boom registered in Myanmar amber. Scientific Reports, 8(1): 16765.

2018. Liu, Z.-J., Wang, X., A novel angiosperm from the Early Cretaceous and its implications on carpel-deriving. Acta Geologica Sinica (English edition), 92(4): 1293-1298.

2018. Santos, A.A., Wang, X., Fu, Q., Diez, J.B., First palynological dating from the Jurassic South Xiangshan Formation (Nanjing area, China). Geobios, 51:559-570.

2018. Wang, X., The Dawn Angiosperms (2nd ed.). Cham: Springer.

2018. Wang, X., An era of errors: Unveiling the truth of Archaeanthus and its implications for angiosperm systematics. ChinaXiv, 201804.201934.

2018. Fu, Q., Diez, J.B., Pole, M., Garcia-Avila, M., Liu, Z.-J., Chu, H., Hou, Y., Yin, P., Zhang, G.-Q., Du, K., Wang, X., An unexpected noncarpellate epigynous flower from the Jurassic of China. eLife, 7:e38827.

2019. Wang, X., Bai, S.-N., Key innovations in transition from homospory to heterospory. Plant Signaling & Behavior 14: 6.

2019. Liu, Z.-J., Hou, Y.-M., Wang, X., Zhangwuia: An enigmatic organ with bennettitalean appearance and enclosed ovules. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 108, 419-428.

2020. Chen, L.-j., Hou, Y.-m., Yin, P.-f., Wang, X., An edible fruit from the Jurassic of China. China Geology, 3(1): 8-15.

2020. Wang, X., Liu, Z.-J., Liu, W., Liao, W., Zhang, X., Liu, Z., Hu, G., Guo, X., Wang, Y.,  Stepping out of the shadow of Goethe: for a more scientific plant systematics. Chinese Bulletin of Botany, 55(4).


Science popularization

2009, Wang, Y, Ma, H, Wang, X., Two interesting stories about silkworms. Evolution of Life, 3:44-45.

2010, Wang, X., On the flower garden in western Liaoning. Life World, 2010, 56-59.



1995, Wang, X. (as a contributor), Atlas of Fossil Plant Anatomy in China, Science Press

2004, Dilcher, D., Lott, T.A., Wang, X., Wang, Q., A history of tree canopies, in Lowman MD, Rinker HB (eds.), Forest Canopies, 2nd Ed. Chapter 6, p118-137, Elsevier Academic Press, Amsterdam

2009, Wang, X., New fossils and new hope for the origin of angiosperms. in: P Pontarotti ed. Evolutionary Biology: concept, modelling and application. Springer-Verlag, Berlin. P51-70.

2010, Wang, X., The dawn angiosperms. Springer, Heidelberg

2010, Cai, C.Y., Wang, X., Fu, Q. (translator), Huxley, T.T. (author), Man’s place in nature and other essays, Beijing, Peking University Press.

2018, Wang, X., The dawn angiosperms. 2nd edition. Springer, Heidelberg.

2018, Wang, X., The dawn angiosperms (in Chinese). Science Press, Beijing.


Symposium Organized

2010, Tracing the Early Angiosperms, 8th European Palaeobotany-Palynology Conference. Budapest.

Community service:
Research Projects:

1999  Grant-in-Aid of Research, Sigma Xi, The Scientific Research Society, Mesofossils, A New Field in Paleobotany, $600

2005  China Postdoctoral Foundation, Study on plant cytoplasm fossils, RMB10,000

2005  State Key Laboratory of Palaeobiology and Stratigraphy, Primary study on plant mesofossils, RMB40,000

2005  Foundation for Talents in Special Specialty, Study on ultrastructures in cytoplasm fossils, RMB50,000

2006  Jiangsu Planned Project for Postdoctoral Research Funds, Study on ultrastructures in cytoplasm fossils, RMB50,000

2006  Royal Society K. C. Wong Fellowship, Cytological study on plant fossils, RMB20,000

2007 Foundation for Talents in Special Specialty, Study on some physiological phenomena in fossil plants, RMB20,000

2007  State Key Laboratory of Palaeobiology and Stratigraphy, Primary study on physiology in fossil plants, RMB20,000

2008 National Natural Science Foundation of China, Study on the ultrastructures and physiological activities in fossil plants, RMB420,000

2008  Scientific Research Foundation for the Returned Overseas Chinese Scholars, State Education Ministry, RMB 25,000

2009  Environment change and life evolution during the Mesozoic in the Upper and Middle Yangtze region, RMB 120,000

2012  Biota changes and their relationships with environment in China in the past 400 million years, RMB 850,000

2013 Team Program of Scientific Innovation and Interdisciplinary Cooperation, Chinese Academy of Sciences, RMB 200,000